Matthew D. Cacopardo
(table available in print form)
s
, number of species
N, number of individuals
R, richness
Ds, Simpson Diversity index
E, evenness
The formulas for R, Ds, and E are as follows:
R =
s
/ log N
Ds = 1 – ( SUM ni (ni – 1)) / ( N ( N – 1))
Dmax = (s-1/s) x (N/N-1)
E = Ds/Dmax